Tropical Animal Behavior

Dr. David Holtzman

July 20-August 14, 2000

Ethology is the study of naturally occurring behavior. In this course, we will examine the rich diversity of animal behavior on the tropical island of Ometepe in Lake Nicaragua. Lectures will cover spatial and foraging ecology, communication, reproductive strategies, and conservation. Both invertebrate and vertebrate species will be used to demonstrate examples of material covered in lectures and to teach ethological methodology. Students

will learn how to form testable hypotheses for field biology and learn different ways of observing, quantifying, and analyzing behavior. This course is intended for advanced undergraduates or beginning graduate students.

Lecture Topics

Ethological approaches and methods
Neural basis of behavior
Orientation, migration and navigation
Spatial learning and memory
Communication
Feeding
Habitat selection
Social behavior
Conservation

Required Text

There is one required text for this course:
Lehner, P. N. (1996) Handbook of Ethological Methods. 2nd edition. Cambridge University Press, Cambridge, UK (ISBN 0-521-63750-3 paperback).

This text can be ordered from your local college bookstore. Please be advised that it may take several weeks for the books to be received. Do not wait until the last minute to order. Please familiarize yourself with the material in this book before you leave for Nicaragua.

Additional Reading

Each student will be responsible for bringing 5 articles relevant to the topics covered in lecture. These articles should concern tropical species and have to be approved by Dr. Holtzman by June 15th, 2000. Each student will be responsible for "presenting" articles to the class. The number of presentations will be dependent on the number of students enrolled. Presentations will involve a houghtful discourse of the hypotheses being asked, the methods used to address those hypotheses, the results of observations or experiments, and the interpretation of those results. In addition, other articles and reference materials will be available to students for preparation of their research reports.

Daily Excursions

The instructors (Dr. Holtzman and graduate teaching assistants) will lead daily trips into the field. Students will be expected to be physically fit and be able to hike continuously for several hours in a day. An overnight excursion will be planned to a cloud forest and crater lake at the top of one of the two volcanoes on Ometepe. These field excursions are intended to demonstrate the complex interactions between animals and their natural environment. Students will learn how to sample different species through the use of standard methods (for example, seining for fish, use of Sherman traps for small mammals, and mist-netting for bats and birds), and methods of ethological observation will also be taught.

A list of necessary equipment and field gear can be found through the La Suerte Field website (http://www.studyabroad.com/lasuerte/addinfo.html#equip). Please be advised that this site lists equipment and materials that are NOT needed for Ometepe. A headlamp and extra batteries are necessary to facilitate hikes after dark. Students should also have a pair of hiking boots (already broken in) and tennis shoes. Because we will be exploring different habitats ranging from lakeshore to tropical lowland forest to mountainous, cloud forest, students should bring appropriate clothing for a variety of environments.

Independent Projects

Each student will form and test a hypothesis about a local species during the last two weeks of the course. The choice of subject species will be left to the student but should be selected from experiences with that species during daily excursions. Each project must also be approved by Dr. Holtzman. The instructors of the course will be available to help guide students design plausible projects, as well as help with the analysis of data collected. Each student will be responsible for writing a 7-10 page research report and presenting their results to the class on the last day of the course.

Pictures from Summer 1999

Further information about this course is available by contacting:

Dr. David Holtzman
Department of Brain and Cognitive Sciences
Meliora Hall
University of Rochester
Rochester NY 14627
Phone: 716-273-4561
FAX: 716-442-9216
E-mail: holtzman@bcs.rochester.edu

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